求分析英语长难句!The introduction of any unmatched transplantation antigens induces the development by the recipient of donor-specific lymphocytes that will produce violent rejection of further transplantations from that donor.这个句子在

求分析英语长难句!The introduction of any unmatched transplantation antigens induces the development by the recipient of donor-specific lymphocytes that will produce violent rejection of further transplantations from that donor.这个句子在
求分析英语长难句!
The introduction of any unmatched transplantation antigens
induces the development by the recipient of donor-specific
lymphocytes that will produce violent rejection of further
transplantations from that donor.
这个句子在"development”后段感觉有歧义啊："of donnor-specific lymphocytes"到底是修饰谁的?感觉没理解清楚
如果是development的,那就是”通过接受者,刺激了一种供者特异性淋巴细胞的发展,这种细胞可以对以后那个供者的移植产生强烈的抵抗作用”
如果是recipient的,则为“通过供者的特异性淋巴细胞的接受者,产生了一种能对以后的移植产生强烈的抵抗作用的发展”
到底哪个对?

求分析英语长难句!The introduction of any unmatched transplantation antigens induces the development by the recipient of donor-specific lymphocytes that will produce violent rejection of further transplantations from that donor.这个句子在
The introduction of any unmatched transplantation antigens induces the development by the recipient of donor-specific lymphocytes that will produce violent rejection of further transplantations from that donor.
that will produce violent rejection of further transplantations from that donor是定语从句,修饰the development.
by the recipient of donor-specific lymphocytes是介词短语,也修饰the development.

根据英文就近修饰的构句特点，应是第二项；其次，根据生物学原理，只有受了某捐献者的移植物，在以后的移植中才能产生更强烈的反应（因为此时抗体产生更快更多）。

The introduction of any unmatched transplantation antigens induces the development by the recipient of donor-specific lymphocytes that will produce violent rejection of further transplantations from t...

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The introduction of any unmatched transplantation antigens induces the development by the recipient of donor-specific lymphocytes that will produce violent rejection of further transplantations from that donor.

of donor-specific lymphocytes是修饰the recipient
然后donor-specific修饰lymphocytes
意思是：受者体内对供体特殊识别的淋巴细胞
development by the recipient of donor-specific lymphocytes
翻译过来就是：
受者体内对供体特殊识别的淋巴细胞分化生长
整句话的意思是：

任何抗原不符移植都会引起受者体内对供体特殊识别的淋巴细胞分化生长，从而导致再次从同一供者移植（组织）时产生强烈的排斥反应。

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